CC Traffic for visibility
It seems we're replying with 501 to requests to swift/varnish upload, where a 416 would be warranted instead, e.g. (I'm posting the logstash id for later reference)
GET https://upload.wikimedia.org/wikipedia/commons/6/6d/Elena_dementieva.ogg bytes=18446744073709550047- _id=AWC2Gg2dKr2U4sZ1omvc
Another source of 501s I've see is specifically from httrack, that seems to be requesting files with range=<last_byte>- (the file below has content-length 6379)
GET https://upload.wikimedia.org/wikipedia/commons/thumb/8/87/Kraków,_Widok_z_wieży_Kościoła_Mariackiego_-_fotopolska.eu_(220013).jpg/120px-Kraków,_Widok_z_wieży_Kościoła_Mariackiego_-_fotopolska.eu_(220013).jpg bytes=6379-
See also swift tests for range requests at https://github.com/openstack/swift/blob/master/test/functional/tests.py#L1087
And the rfc for range requests for reference https://tools.ietf.org/html/rfc7233