- Using Firefox 57 or Chrome 61 (e.g. via BrowserStack), released in 2017.
- Log in via https://en.wikipedia.org to a simple account with a username and password (i.e. no 2FA, WebAuthn, autoconfirmed status, skipcaptcha right, bot right, staff group, etc; could be ten years old or recently created). In my case KrinkleSock on enwiki.
Actual
WebAuthn is interferring with the login of an account that does not use WebAuthn, and is turning its (optional) JavaScript payload for Grade A payload into a mandatory and thus impossible challenge for Grade C browsers.
JavaScript must be enabled in your browser to use a security key.
Other information
"oathauth-webauthn-javascript-required": "JavaScript must be enabled in your browser to use a security key.",

