Split from T195768.
Start translating an article with an audio template - e.g. enwiki 'Meow'. All links inside the audio template will be displayed red in the source article:
vs the original article:
The issue is not limited to the beginning of an article - if the audio template is in the middle (e.g. https://en.wikipedia.org/wiki/Black-capped_chickadee), the issue is still present: